Quote:
When the lifter begins to roll up the cam lobe, the ball check is held against its seat in the plunger by the ball check spring, which traps the oil in the base of the lifter body below the plunger. The plunger and lifter body then raise as a unit, opening the valve. The force of the valve spring, which is exerted on the plunger through the rocker arm and pushrod causes a slight amount of leakage between the plunger and the lifter body.
This leak-down allows a slow escape of trapped oil in the base of the lifter body. As the lifter rolls down the other side of the cam lobe and reaches the base circle or "valve closed" position, the plunger spring quickly moves the plunger back to its original position. This movement causes the ball check to open against the ball spring, and oil from within the plunger is drawn into the base of the lifter. This restores the lifter to zero lash.
Long story short... a "zero lash" valve clearance (in this instance) also means zero preload... and it can compensate (within a narrow range) for variations in head gasket thickness or block deck/head deck dimensional variations.